This is going off of the voice lines used during the Archon Hunt mission, so look there real quick if you want some context: Link
Ch 1: The Archons
According to Pazuul and the Lotus, the Archons were originally Sentients created by Hunhow and Praghasa, modeled after animals of Earth; Amar, the Wolf. Boreal, the Owl. Nira, the Snake. And Pazuul, the Ram. Together, they helped rule over the rest of the Sentients during their days on Tau.
I guess they may have fought in the Old War, and died during it. This would explain how they are back during the events of the New War; Their bodies were recovered and hybridized with the corpses of dead Warframes in order to bring them back to life. However, the Archons did not like this, and it filled them with hatred. "Boreal is consumed by hatred. He blames both your people and ours for what was done to him.- "To be mutilated like this, turned into a weapon, was humiliating for him." "Archon Amar yearns for the new creation that Pazuul has promised him. He imagines he will be a lord of the wilderness, instead of what he is now. -" - Lotus.
That would make sense. You die in a war, only to be brought back to life by your own people and find out that you're grafted to the body of your worst enemy, which is the only thing keeping you alive. You are then sent out as a tool to fight and die again.
I think Boreal is the only one that is deeply affected by this, as he hates both the Tenno and the Sentients for what happened to him. The Tenno were the ones who killed him during the Old War, while the Sentients, specifically Erra, brought him back to life to be used as a weapon.
Ch 2: The Creation of Pazuul
During the events of the New War, we see Erra get crushed by falling debris when helping us fight the final Archon, and presume him dead. We then see him again, with the head of the Ram sitting on his shoulders. Erra does seem to be in there somewhere, as his voice does regress back during some of his lines. "Tenno... I'm truly sorry... for everything. I saw the truth far too late. She will never forgive me. But you will... won't you?" - Erra
So it could be said that he may not have died during the War, and had the head of Pazuul grafted onto him when he was injured.
I'm not exactly sure who did this to him. One of Pazuul's voice lines mentions his mother, Praghasa. "Bite my head clean off, mother. That's where the good meat is. Suck the guts until the little legs kick, kick, kick." - Pazuul. Praghasa was the one who created the Sentients, acting similar to a queen bee. So maybe she was the one who did this to Erra; She 'bit his head off' and replaced it with the head of Pazuul. I'm not sure why she would do that though.
According to voice lines from the Lotus, Archon Nira may also have something to do with it. "Nira. Could she somehow be responsible for what Erra has become? A repayment for what he did to her? The Serpent is old, Tenno, and full of secrets." - Lotus. As I said earlier, the Beasts hated being turned into Archons. Nira may have had Erra converted into an Archon as revenge for turning her into one. She also may be the one who keeps Erra's personality suppressed under Pazuul's. "Who whispered dreams of dominion into Pazuul's ear? Who keeps Erra suppressed and tormented with dilusions? The stench of Nira's venom is upon this, Tenno." - Lotus. This would also relate to what Lotus says about not knowing if Pazuul is the Archon's leader, or puppet. They could have had Erra converted as revenge, intending to use him as a tool to go back to Tau.
Ch 3: Pazuul's Plan
During the Archon Hunts, we're told by Lotus that Pazuul is building something. We're not given a hint as to what, except that it's taking a lot of power and resources to be constructed, and that both Narmer and the Archons are waiting for it. I think it may have something to do with Pazuul's goal of going back to Tau. Although, I don't know how he intends to get there. If he wanted to, he could probably do the same thing Ballas did, and use Praghasa to eat the sun again. However, that wouldn't match up with him making something. I think that it's either a Void portal, or a Solar Rail that leads to Tau.
That's kinda all I got. Not sure how to end this, though. I gotta say, though, the Archons might be my favorite characters in the lore now. There's a lot going on when you think about it.
Thanks for reading.
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source https://www.reddit.com/r/Warframe/comments/xdl3p4/who_are_the_sentient_beasts_and_what_is_pazuul/
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